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Speothos

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Genus of carnivores

Speothos
Temporal range: Pleistocene to Recent
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Canidae
Subfamily: Caninae
Tribe: Canini
Genus: Speothos
Lund, 1839
Type species
+Speothos pacivorus
Species
Synonyms
  • Icticyon Lund, 1843
  • Abathmodon Lund, 1843
Speothos venaticus

Speothos is a genus of canid found in Central and South America. The genus includes the living bush dog, Speothos venaticus, and an extinct Pleistocene species, Speothos pacivorus. Unusually, the fossil species was identified and named before the extant species was discovered, with the result that the type species of Speothos is S. pacivorus. S. pacivorus had a larger overall body size and a double-rooted second lower molar. It has been proposed that Speothos originated in the Brazilian highlands sometime during the Pleistocene.[1]

The paleobiogeography and evolutionary relationships of Speothos and other South American canids is unclear. Morphological data suggest that Speothos is most closely related to another small canid, Atelocynus (short eared dog). The skulls of both Speothos and Atelocynus have short nasals that terminate rostral to the maxillary-frontal suture, a character shared with Vulpes (fox genus). Speothos and Atelocynus also possess very small frontal sinuses that are minimally expanded. Recent analyses of molecular (mtDNA) data alone and in combination with morphological data support a sister-taxon relationship between Speothos and Chrysocyon (maned wolf); it has been suggested that these highly derived canids diverged at least 3 million years ago, invading South America as separate lineages.

Speothos is distinctive in that it shares similar hypercarnivorous modifications of the dentition with Cuon (dhole) and Lycaon (African wild dog). The crushing role of the post-carnassial molars is reduced. The lower carnassial (m1) of Speothos lacks the entoconid, creating a trenchant (cutting) heel on the tooth. The hypoconid on the heel (talonid) of the m1 fits into a basin in the M1 when the teeth are in occlusion. Speothos and Cuon exhibit reduction in both number and size of the post-carnassial molars.[2]

References

[edit]
  1. ^ Berta, Annalisa (30 November 1984). "The Pleistocene Bush Dog Speothos pacivorus (Canidae) from the Lagoa Santa Caves, Brazil". Journal of Mammalogy. 65 (4): 549-559. doi:10.2307/1380837. JSTOR 1380837.
  2. ^ Owens, Pamela (2001). "Speothos venaticus (South American bush dog)". Digital Morphology. Retrieved 2022-03-02.


Extant Carnivora species
Prionodon (Asiatic linsangs)
Pantherinae
Neofelis
Panthera
Felinae sensu stricto
Bay cat
lineage
Pardofelis
Catopuma
Caracal
lineage
Caracal
Leopardus
Lynx
Puma
lineage
Acinonyx
Puma
Leopard cat
lineage
Prionailurus
Felis
Viverroidea
    • see below|
Hemigalinae
Paradoxurinae
Paradoxurus
Viverrinae sensu lato
Viverrinae
sensu stricto
Viverra
Poiana
(African linsangs)
Genetta
(genets)
Herpestoidea
    • see below|
Hyaenidae
(hyenas)
Proteles
Hyaeninae
(bone-crushing hyenas)
Crocuta
Herpestidae sensu lato
Eupleridae
(Malagasy
carnivorans)
Euplerinae
(Malagasy civets)
Eupleres (falanoucs)
Galidiinae
(vontsira)
Galidictis
Salanoia
Herpestidae sensu stricto (mongooses)
Suricata
Mungos
Helogale
Crossarchus
(kusimanses)
Urva
(Asian mongooses)
Bdeogale
Herpestes
(slender mongooses)
Urocyon
Nyctereutes
(raccoon dogs)
Vulpes
(true foxes)
Speothos
Lycalopex
(South American foxes)
Lupulella
Lycaon
Canis
Ailuropoda
Tremarctos
Ursinae
Ursus
Mustelida
Pinnipedia (seals)
    • see below|
Musteloidea
    • see below|
Odobenidae
Callorhinus
(northern fur seals)
Otariinae
(sea lions)
Zalophus
Neophoca
Arctocephalus
(southern fur seals)
Phoca
Pusa
Monachini
(monk seals)
Neomonachus
Mirounga
(elephant seals)
Lobodontini
(Antarctic seals)
Ailuridae
Conepatus
(hog-nosed skunks)
Mephitis
Mydaus
(stink badgers)
Spilogale
(spotted skunks)
Bassariscus
Procyon
(raccoons)
Bassaricyon
(olingos)
Nasuina
(coatis)
Nasua
Nasuella
(mountain coatis)
Mustelidae
    • see below|
Mellivora
Arctonyx
(hog badgers)
Meles
(Eurasian badgers)
Melogale
(ferret-badgers)
Pekania
Gulo
Martes
(martens)
Lyncodontini
Galictis
(grisons)
Ictonychini
(African polecats)
Vormela
Ictonyx
Lontra
Enhydra
Lutra
Lutrogale
Aonyx
Neogale
(New World weasels)
subgenus Mustela
(paraphyletic)
subgenus Lutreola
(paraphyletic)
subgenus Putorius
Extinct Canidae
+Hesperocyoninae
Osbornodon
Phlaocyonini
Phlaocyon
Cynarctina
Aelurodontina
Borophagina
Borophagus
Caninae
    • see below|

Urocyon
Nyctereutes
(raccoon dogs)
Vulpes
(true foxes)
Cerdocyonina
(zorro)
Speothos
+Dusicyon
Lycalopex
(South American foxes)
Canina (wolf-like canids)
    • see below|

+Eucyon
Lycaon
Dhole (Cuon alpinus)
Coyote (C. latrans)
Red wolf (C. rufus)
+Prehistoric
+Recently
extinct


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