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Thessaly and surrounding regions
The political history of the other regions of Greece during this period is no less complex. Thessaly was ruled in its eastern parts by the Franks after 1204, while the western regions were disputed by the rulers of Epirus and Nicaea. About 1267 John I Doukas established himself as independent ruler, with the Byzantine title sebastokrator, at Neopatras, but in expanding his control eastward he came into conflict with Michael VIII, whose attacks he repelled with the assistance of the dukes of Athens and Charles I of Anjou. Venetian support, the result of a favourable trading relationship (Thessaly exported agricultural produce), helped maintain Thessalian independence until the arrival in 1309 of the Catalan Grand Company. This band of Spanish mercenaries, who originally had been hired by Andronicus II to fight the Seljuqs in Anatolia, had turned against imperial authority and established themselves in the Gallipoli peninsula. From there they moved into Greece through Thrace and Macedonia, which they plundered, and from 1318 onward they occupied the southern districts of Thessaly. The northern regions remained independent under the ruler Stephen Gabrielopoulos until 1332, and then they were taken by John II Orsini of Epirus. In 1335 Thessaly was retaken by the Byzantine Empire, and from 1348 it acknowledged the overlordship of the Serbian ruler Stefan Dusan. After his death (1355) the self-styled emperor Symeon Uros, despotes of Epirus and Akarnania, was able to seize control of both Epirus and Thessaly and rule independently following the death of Nikephoros II in 1358/59. He was succeeded by his son John, who adopted the monastic life in 1373. The caesar Alexios Angelos Philanthropenos took control, governing as a vassal of the Byzantine emperor John V, but in 1393 the conquest of Thessaly by Ottoman forces put an end to its independence.
Athens, Thebes, and Corinth
In the south Greece was divided among a number of competing political units. After 1204 the dukes of Athens (mostly of French or Italian origin) controlled much of central Greece, with their main base at Thebes. They had political interests to the north and in the Peloponnese. However, in 1311 the Catalan Grand Company established its power over the duchies of Athens and Thebes, turning out their Latin lords. Under the protection of the Aragonese king Frederick II of Sicily (three sons of whom became dukes of Athens), they dominated the region until the Navarrese Company (an army of mercenaries originally hired by Luis of Evreux, brother of Charles II of Navarre, to help assert his claim over Albania and then temporarily in the service of the Hospitallers, a military-monastic order) took Thebes in 1378 or 1379. This weakened Catalan power and opened the way for the Florentine Acciajuoli, lords of Corinth, to take Athens in 1388. The latter then ruled all three regions until their defeat at the hands of the Ottomans in the 1450s.
The Peloponnese
In the Peloponnese the political rivalry between the Byzantines and the Frankish principality of Achaea dominated. The principality was at its most successful under its prince William II Villehardouin (1246-78), but in 1259 he had to cede a number of fortresses, including Mistra, Monemvasia, and Maina, to the Byzantines. Internecine squabbles weakened resistance to Byzantine pressure, especially from the 1370s onward, when Jacques de Baux hired the Navarrese Company to fight for his claim to the principality. Upon his death in 1383, the Navarrese exercised effective political control over the Frankish territories under the commanders of the company. The last Navarrese prince, Pierre de Saint-Superan, joined the Ottomans in 1401 to raid Byzantine possessions in the southern Peloponnese; he died in 1402. He was succeeded by his widow, Maria Zaccaria, representative of an important Genoese merchant and naval family. She passed the title to her nephew Centurione II Zaccaria, who lost much of the territory to the Byzantine despotate of the Morea. In 1430 he married his daughter to the Byzantine despotes Thomas Palaeologus, handing over his remaining lands as her dowry. From this time on, the Byzantine despotate of the Morea effectively controlled most of the Peloponnese. However, the Ottoman presence and the fall of Constantinople to Sultan Mehmed II in 1453 effectively ended this final period of Byzantine rule. The Morea resisted Ottoman conquest until 1460, when it was finally incorporated into the Ottoman Empire (a year earlier than the empire of Trebizond, which fell in 1461). All of Greece was by this time under Ottoman authority, with the exception of some of the islands, which retained a tenuous independence under Venetian or Genoese protection.
Serbian and Ottoman advances
Byzantine power in the northern Greek regions was effectively destroyed by the expansion of the Serbian empire under Stefan Dusan, the results of which included the loss of Epirus, Thessaly, and eastern Macedonia. From the 1350s the Ottomans established themselves in Europe, taking the chief towns of Thrace in the 1360s and Thessalonica in 1387. Apart from the Despotate of the Morea, therefore, and certain of the Aegean isles, there remained in Greece no Byzantine imperial possessions by the beginning of the 15th century.
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